These are particularly evident in the sexually dimorphic phenotypic traits that distinguish the sexes of a species, but unlike the sex organs, are not directly part of the reproductive system.
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In humans, visible secondary sex characteristics include pubic hair, enlarged breasts and widened hips of females, and facial hair and Adam's apple on males.
Biologists today distinguish between "male-to-male combat" and "mate choice", usually female choice of male mates.
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Sexual differentiation begins during gestation, when the gonads are formed.